Contributions to Zoology, 86 (1) – 2017Isabel T Hyman; Irantzu de la Iglesia Lamborena; Frank Köhler: Molecular phylogenetics and systematic revision of the south-eastern Australian Helicarionidae (Gastropoda, Stylommatophora)

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Cucullarion Stanisic, 1998

Cucullarion Stanisic, 1998: 597.

Type species: Cucullarion parkini Stanisic, 1998, by original designation; masculine


Diagnosis. Shell. Medium sized, reduced, ear-shaped, flattened; base and internal whorls completely membraneous, dorsal calcified area plate-like; shell glossy, no sculpture.

Animal. Beige to orange- or red-brown; caudal horn small. Mantle lobes large, left and median lobes fused to form cephalic shield; shell lappets large, fused at both sides, covering most of shell (Fig. 32A-B).


Fig. 32. Live specimens of Cucullarion, Peloparion and Ubiquitarion gen. nov. A. Cucullarion parkini, AM C.512610, Lamington National Park. B. C. albimaculosum, AM C.103209 (photographer: G. Millen). C. Peloparion helenae, MV F219249, Barrington Tops National Park (photographer: A. Moussalli). D. Ubiquitarion iridis, AM C.512394, Pikapene National Park. Scale bars = approx. 10 mm.

Gentitalia. Ovotestis of 3-4 lobes grouped together and embedded in tip of digestive gland. Talon and carrefour embedded in albumen gland. Spermoviduct folded 5 times. Free oviduct long with no internal sculpture; capsular gland small, spherical. Bursa copulatrix moderately long, reaching two thirds of the way along spermoviduct, consisting of swollen base, slender duct and elongate sac; internally with longitudinal pilasters; inserted on vagina. Vagina very short. Penial tunica tightly attached to epiphallus by muscle fibres; epiphallus enters penis through verge; epiphallic caecum absent; epiphallic flagellum with axial filament present; single internal crypt present in part of epiphallus adjacent to flagellum but not in flagellum. Spermatophore a soft-walled capsule with hard tail-pipe; one spine present at junction of capsule and tail-pipe; tail-pipe sculptured with a spiraling row of tiny teeth.

Remarks. Cucullarion was introduced for a semislug from southern Queensland, C. parkini (Stanisic, 1998) and was later expanded to include a second species, C. albimaculosum (Stanisic et al., 2010). The two species are notable for their degenerate, plate-like shells with membranous sides, bases and internal whorls.