Contributions to Zoology, 86 (1) – 2017Isabel T Hyman; Irantzu de la Iglesia Lamborena; Frank Köhler: Molecular phylogenetics and systematic revision of the south-eastern Australian Helicarionidae (Gastropoda, Stylommatophora)

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Ubiquitarion gen. nov.

Type species: Peloparion iridis Hyman, 2007

Description. Shell. Small, ear-shaped, flattened, last whorl large, underside membranous, umbilicus closed, shell glossy, protoconch and teleoconch with no visible sculpture (Table 3, Fig. 35B).

Animal. Beige with yellowish and black markings and iridescent specks of pigment. Mantle lobes and shell lappets well developed.

Genitalia. Ovotestis of one lobe, embedded near apex of digestive gland. Talon and carrefour embedded in albumen gland. Spermoviduct folded three times. Distal portion of free oviduct with small spherical capsular gland with no internal sculpture, remainder of free oviduct with longitudinal pilasters internally. Bursa copulatrix moderately long, reaching halfway along spermoviduct; internally with longitudinal pilasters (duct only); inserted on vagina. Vagina short, internally with longitudinal pilasters. Epiphallus enters penis through thickened ring; epiphallic caecum absent; descending arm of epiphallus (near flagellum) encased in a thick muscular sheath; epiphallic flagellum with axial filament present, also encased in thick muscular sheath. Spermatophore a soft-walled capsule with hard tail-pipe; one branching spine present on capsule; double row of branching spines present in spiraling pattern along tail-pipe.

Remarks. This genus is created for Peloparion iridis. This species is superficially similar to Peloparion helenae, but differs anatomically, particularly in the penial complex and spermatophore. Mitochondrial DNA sequences also show that the two species do not group together.

Etymology. Combination of ubique (Latin = ‘everywhere’; arion (Latin) referring to a ‘kind of snail or slug’; masculine.