Contributions to Zoology, 77 (4) - 2008Marco A. Bologna; Andrea Di Giulio: Revision of the genus Trichomeloe Reitter, with the description of new species and fi rst instarlarvae (Coleoptera: Meloidae)
Appendix

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Appendix I. Annotated catalogue and descriptions of two new species

For each species are indicated: the synonymies and the main citations from the literature; the type locality and the type material, also for synonyms; a short diagnosis and some taxonomical remarks; the list of localities with the acronym of collection and/or literature citations, divided in countries, and also provinces in the case of Turkey, the larger state. Two new species from the Iraki Mesopotamia and Syria respectively are also described.

Genus Trichomeloe Reitter, 1911

Meloë (Trichomeloë) Reitter, 1911: 391; Cros, 1934: 431

Trichomeloe, MacSwain, 1956: 71 (pars); Bologna, 1983: 24 (pars); Bologna, 1989: 359; Bologna, 1991: 109; Bologna and Pinto, 2002: 2055.

Type species: Meloe chrysocomus Miller, 1861: 206, by subsequent designation (MacSwain, 1956).

a) Group of Trichomeloe chysocomus

Trichomeloe chrysocomus (Miller, 1861) (Figs 6a, 8a)

Meloë chrysocomus Miller, 1861: 206; Escherich, 1890: 88, 94.

Meloë (Trichomeloe) chrysocomus, Reitter, 1911: 391.

Trichomeloe chrysocomus, MacSwan, 1956: 71; Bologna, 1988: 30; Bologna, 1989: 359; Bologna, 1991: 110

Type material. Types of T. chrysocomus were not examined; they are probably preserved in the Motschulsky’s collection, which is divided between the Moscow State University, the Zoological Museum of Saint Petersburg, the Humboldt Museum and the German Entomological Institute. The type locality is “Syrie” (Miller, 1861).

Diagnosis. Integument black, subopaque, with one small frontal red spot; setation dense both dorsally and ventrally, setae light golden-yellow, short, recumbent, longer on head, pronotum and elytra, producing a brownish shade to the body. Body length: 11.8-20 mm. Head subequal or scarcely wider than pronotum, eye small, slightly convex; templa 2.5 as long as the longitudinal diameter of eye, sub-parallel, slightly wider and rounded posteriorly; middle frontal line superficial; punctures fine, shallow, distanced, intermediate surface shagreened. Antennomeres elongate and subcylindrical; I elongate and widened at apex, II ca 0.5 as long as III subglobose, III-V progressively shorter, VI-X subequal and similar in length to V, XI twice as long as X, parallel and conically narrowed at apex; male segments III-VII unmodified. Pronotum reniform, almost twice as wide as long, sides anteriorly evidently curved; base depressed, middle surface longitudinally slightly depressed; punctures as on head. Elytra vaguely rugulose. Male genitalia (Fig. 6a): in dorsal view, gonocoxal piece short, slightly wide in the middle, gonostyli conically narrowed with stout apical lobes; in lateral view, gonostyli narrow, apical lobe evidently curved and stout, microsetose; aedeagus in lateral view with one single apical hook on ventral side, short and minute.

Remarks. Among numerous specimens from Hasanbeyli, southern Turkey, few of them have a darker setation, rather than light.

Distribution (Fig. 8a, triangle). CYPRUS: Cyprus (MRSN; Baudi, 1878; Cros, 1934; Bologna and Marangoni, 1990: 354).

TURKEY: (ADANA) Bahçe, Hasambeyli (CB). (HATAY) Akbès (HNHM as deflexus; JP; MNHN). (MARDIN) Akresta gecidi (CK). (ADYAMAN) Nemrut Daği (CSa). (SANLI URFA) near Halfeti, (CB); 15 km S Hilvan (CB).

SYRIA: Syria (HNHM; MNHN; NHMW; Miller, 1861; Baudi, 1878; Escherich, 1890; Reitter, 1895; Pliginskji, 1914; Cros, 1934; Chikutanov, 1999); Halep (CB; JP; HNHM; MHNG; MNHN); Halep, Tal Hadia (HNHM); Tell Halaf, Guzana, Djirdjibufer steppe (MNHU); 48 Km NW Hama, Ruins of Afamiya (Afamea) (CB); Jeirud (CW).

ISRAEL and PALESTINE: Israel (Chikutanov, 1999); Palestine (Cros, 1934); Jerusalem (MSNV; MCSN; MNHN; Pic, 1900); near Jerusalem, Chan Hatrura (Shalberg, 1913); Nazreth (Sahlberg, 1913); Kana (Sahlberg, 1913); Jordan River (Sahlberg, 1913); Guesthouse between Garimherzig and Samariter (MNHU). Uncertain localities: Moab or Moag or O’Megib? (MNHN); Lenal ? (HNHM). Chikatunov (1999) cited the species from the following areas: Upper Galilee, Valley of Yzre’el, Judean Hills, southern coastal plain, Dead Sea, northern Negev, central Negev. An undetermined species of Trichomeloe, possibly referable to this species, is cited by Finkel et al. (2002) from the Lower nahal Keziv, in the Western Upper Galilee, Israel.

JORDAN: Kerak, Wadi Al Hasa (CB).

Trichomeloe conicicollis (Reitter, 1907) (Figs 5a, 5c, 6b, 8a)

Meloë conicicollis Reitter, 1907: 215; Kaszab, 1968: 447.

Meloë (Trichomeloe) conicicollis, Reitter, 1911: 391.

Trichomeloe conicicollis, Bologna, 1988: 30; Bologna, 1989: 359; Bologna, 1991: 110.

Type material. In the HNHM is preserved the holotype, we examined, labelled “Monotypus 1907 Meloe conicollis Reitter” (white, with red frame, not orginal), “collez. Reitter” , “Adana” (handwritten, white) (see Reitter, 1907).

Diagnosis. Integument shiny black with one small frontal red spot; setation sparse dorsally and denser ventrally, setae dorsally black-greysh and ventrally brownish, recumbent, short particularly on head, pronotum and elytra. Body length: 11.9-16 mm. of body. Head elongate, posteriorly obtusely subtrapezoidal, scarcely wider than pronotum, eye wide, convex; templa about 2.5 as long as the longitudinal diameter of eye, parallel, posteriorly elongate and converging on the occiput which appears almost potruded; middle frontal line superficial or well visible; punctures uniformly wide, deep and relatively dense, intermediate surface shagreened. Antennomeres (Fig. 5a) elongate and subcylindrical; I elongate and widened at apex, II slightly less than 0.5 than III, subglobose, III about 1.5 as long as the followings, IV-X subequal and similar in length, XI about twice as long as X, parallel and conically narrowed at apex; male segments III-VII unmodified. Pronotum (Fig. 5c) elongate subcampaniform, at base narrower than elytra, wider than long at base, sides progressively oblique and anteriorly curved; base depressed, middle surface longitudinally slightly depressed, sides vaguely depressed; punctures as on head, but in some specimens rugosities are also present, particularly at base. Elytra with longitudinal, superficial rugosities. Protarsi and mesotarsi of male very short. Abdominal tergites deeply punctured. Male genitalia (Fig. 6b): in dorsal view, gonocoxal piece quite elongate, gonostyli setose and slightly depressed at base, conically narrowed with slender apical lobes, with long setae on both sides; in lateral view, gonostyli very narrow and slender, progressively producing in an apical lobe, slightly curved and with elongate setae; aedeagus in lateral view with one single apical hook on ventral side, short and minute, oblique and acute.

FIG2

Fig. 5. Trichomeloe conicicollis, male: a) left antenna, dorsal view; c) pronotum. T. sericellus, male: b) left antenna, dorsal view; d) pronotum. T. deflexus, male: e) prosternum; f) mesosternum; g) last visible abdominal sternite. Scale bars = 1 mm.

Remarks. This species differs greatly from others by the shape of occiput, pronotum, and the elytral rugosities. Also the shape of male genitalia greatly distinguish T. conicicollis from all other species, T. chrysocomus included.

Distribution (Fig. 8a, square). GREECE: Dodekanissa, Castelorizo Island (MSNM; MSNV; Schatzmayr, 1941; Bologna, 1986; Bologna and Marangoni, 1990: 353; Bologna, 1994: 10).

TURKEY: (MUĞLA) (CB); Fethye, Dodurga (NHMW; Kaszab, 1968). (ANTALYA) Lycian Taurus (HNHM; MNHU; MZUA); Kaş (CB; MRSN). (ADANA) Adana (HNHM; Reitter, 1907). Other specimens with no detailed localities (HNHM; NHMW).

(b) Group of Trichomeloe sericellus

Subgroup A

Trichomeloe deflexus (Reitter, 1889) (Figs 5e-5g, 6c, 8b)

Meloë deflexus Reitter, 1889: 106; Escherich, 1890: 88, 95; Reitter, 1895: 13.

Meloë (Trichomeloe) deflexus, Reitter, 1911: 391.

Trichomeloe deflexus: Bologna, 1988: 30; Bologna, 1989: 359; Bologna, 1991: 110.

Type material. Reitter (1889) indicated “Syrien” as Type locality, but this must be actually restricted to “Jerusalem”. In fact, in the HNHM are preserved the following specimens we examined: holotype, labelled “ Meloe deflexus Reitter”(white, red framed), “coll. Reitter” (red on white), “Jerusalem” (printed, white), “U. Sahlb.” (printed, white); 1 paratype male, with the same labels of the holotype; 1 paratype male, with similar labels, actually belongs to T. sericellus.

Diagnosis. Integument leaden black, subopaque, with one small red frontal spot widened anteriorly, subtriangular or oval; setation short, quite dense both dorsally and ventrally, setae light yellow-grey, short, longer on head and pronotum where it is recumbent along the depressions. Body length: 10-18.6 mm. Head subequal or scarcely wider than pronotum, eye small and convex, in lateral view fore margin slightly incised, short and only slightly narrowed anteriorly; templa 2.5 as long as the longitudinal diameter of eye, sub-parallel, slightly wider and rounded posteriorly; middle frontal line superficial; punctures on head and pronotum deep and denser, intermediate surface shagreened. Antennomeres elongate and subcylindrical; I elongate and widened at apex, II ca 0.5 as long as III subglobose, III antennomere as long as IV; VI-X subequal and similar in length to V, XI twice as long as X, parallel and conically narrowed at apex; male segments III-VII modified to form an anterior almost impunctate, shagreened area, extended also dorsally, III-VI in lateral vision not distinctly widened at apex. Pronotum subreniform, almost twice as wide as long, sides laterally parallel and anteriorly slightly curved; base depressed, middle surface longitudinally slightly depressed; punctures as on head. Elytra obsolescently rugulose. Male genitalia (Fig. 6c): in dorsal view, gonocoxal piece short, wide in the middle, gonostyli conically narrowed with stout apical lobes; in lateral view, gonostyli narrow, apical lobe slender, elongate, with long setae on both sides; aedeagus in lateral view with two apical hooks on ventral side, both short, the proximal one minute.

FIG2

Fig. 6. Male genitalia in ventral and lateral view and aedeagus in lateral view of: a) Trichomeloe chrysocomus; b) T. conicicollis; c) T. deflexus; d) T. syriacus; e) T. mesopotamicus; f) T. sericellus; g) T. ottomanus. Scale bars = 1 mm.

Distribution (Fig. 8b, spot). TURKEY: (ADIYAMAN) Narince (Özbek and Szaloki, 1998); Karababa bridge (Özbek and Szaloki, 1998). (HATAY) Tapbogazi, Issuspass, Amanus (Kaszab, 1968; Özbek and Szaloki, 1998); Akbès (MNHN). (GAZIANTEP) Sakcagöz (NHMW; Kaszab, 1968); Yavuzeli NW of Gaziantep (NHMW, Kaszab, 1968); (KAHRAMAN MARAS) Pazarcik (MSNV).

SYRIA: Syria (HNHM; MNHN; NHMW; Reitter, 1889; 1895; Escherich, 1889; 1890 Özbek and Szaloki, 1998); Damaskus (HNHM; MNHN); Basalt Rock 6 km N of Salhaa (HNHM); Ganawat (CW); Es Suweida (CW).

ISRAEL and PALESTINE: Palestine (HNHM); Jaffa (NHMW); Nablous (MNHN); Jerusalem (BMNH; HNHM; JP; MCSN; MNHN; MSNV; MSNM; NHMW; Sahlberg, 1913); Jerusalem Mt. Sion (MNHN); Jerusalem, Mount of Olives (Sahlberg, 1903); between Jericho and Jerusalem (MNHU); Ghor (MNHN); Jordan Valley, S Mehola, Wadi Malih (CB; SMNS); Bethlem Eastern side (HNHM); Maale, Adumim (CB); Emek Isaac (HNHM); Beer Sheba (CB); Beer Sheba near Tümpel (NHMW); NE Beer Sheba (CB; NHMW).

JORDAN: Jerash (CB); Zerka (MHNG); E Jordan, Wadi Schaib (MHNG); E Jordan, Fuhes (MHNG); S Jordan, Wadi Waba S Makba (HNHM); S Jordan, Madaba (MHNG); S Jordan, Wadi Wala, S Madaba (MHNG); Abu Nusayr, 32°05’N-35°52’E; Al Azraq 31°52’N-36°50’E; Al Jubayhah 32°01’N-35°52’E; Al Karak 31°11’N-35°42’E; Al Kufrayn 31°50’N-35°55’E; Al Muqabalayn 31°54’N-35°54’E; Amman 31°57’N-35°56’E; As Salt 32°03’N-35°44’E; Gawr Kabed 32°04’N-35°34’E; Irbid 32°33’N-35°51’E; Khilda 32°00’N-35°51’E; Na’ur 31°53’N-35°50’E; Sahab 31°53’N-36°00’E; Suwaylih 32°02’N-35°50’E; Tabarbawr 32°00’N-35°57’E; Zavy 32°06’N-35°43’E (Katbeh-Bader, 1996: some of these records could refer to other species).

Trichomeloe mesopotamicus Bologna n. sp. (Figs 6e, 8b)

Type material. 1 male holotype and 2 males and 2 females paratypes, “Mesopotamia Mosul” (MNHU, identified as chrysocomus, 1 CB); 1 male paratype, “Mesopotam. Assur, 08”; (NHMW, identified as “chrysocomus Dr. Z. Kaszab, 1959”); 2 females paratypes, “Mesopotamia Mosul, coll. Hauser v.09” (NHMW); 1 male paratype, “Assur Mesopotam. Pietschmann .10”; “Mesopotam. Esp. Nat. Ö.V. 1910”; chrysocomus Miller det. Breit” (NHMW); 1 female paratype “Mesopot. Assur 08”; (NHMW, identified as “Meloe deflexus Dr. Z. Kaszab 1959”); 1 male paratype, “Mesopotamia Mossul coll. Hauser 5.09” (NHMW); 1 female paratype, “Assur Mesopotam. Pietschmann .10”; “Mesopotam. Esp. Nat. Ö.V. 1910” (NHMW).

The type locality, Mosul, is located in the Northern Irak; the locality of Assur is located on the right side of the Tigris River, about 110 km South of Mosul.

One male paratype (CB), lacks V-XI right antennomeres, and another male paratype (NHMW) lacks right metatarsomeres III-IV.

Other specimens examined: 1 male Mosul (HNHM), 1 male Assur (IV.1908 HNHM).

Description. Integument black, opaque, with one small transverse frontal red spot; setation very dense both dorsally and ventrally, setae light golden-yellow, producing a brownish shade to the body, elongate on the whole body, on the pronotum slightly recumbent along the depressions, slightly shorter on elytra and ventrally. Body length: 10-16.2 mm.

Head subequal or scarcely wider than pronotum, eye dorsally only slightly convex, in lateral view fore margin incised, narrowed anteriorly; templa 2.5 as long as the longitudinal diameter of eye, subparallel, slightly wider and rounded posteriorly; middle frontal line superficial; punctures on head and pronotum slightly deep and distanced, intermediate surface almost umpunctured. Antennomeres elongate and subcylindrical; I elongate and widened at apex, II ca 0.5 as long as III, subglobose, III shorter than IV, VI-X subequal and similar in length to V, XI twice as long as X, parallel and conically narrowed at apex; male segments III-VII modified to form a wide anterior almost impunctate, shagreened area, slightly narrower on VII.

Pronotum subreniform, almost twice as wide as long, sides more obliquely narrowed anteriorly where are slightly curved; base depressed, middle surface longitudinally depressed; punctures as on head. Elytra vaguely rugulose.

Male genitalia (Fig. 6e): in dorsal view, gonocoxal piece slightly elongate, wider anteriorly, gonostyli quite short and with stout apical lobes; in lateral view, gonocoxal piece narrow and elongate, gonostyli subcylindrical on the basal third, then dorsally curved in the middle and suddenly inclined to form a short apical lobe, with very long dorsal setae on the apical third, almost straight on ventral margin; aedeagus in lateral view with two apical hooks on ventral side, both short.

Remarks. This species is phenetically very similar to T. chrysocomus based on integument and setation colour, and the pronotum shape; some paratypes were previously identified as chrysocomus by the late specialist Dr. Z. Kaszab. It actually belongs to another group of species, based on the modified male antennae consisting of a distinct raised, nude and smooth area on segments III-VII, and the aedeagus with two hooks.

Among the group of species related to T. sericellus, T. mesopotamicus appears more similar to T. deflexus because of the slender antennomeres, head colouration, and the sub-depressed pronotum. The main differences characterizing T. mesopotamicus vs. T. deflexus include: integument black-brown more subopaque; red frontal spot transverse vs. elongate in front; setation rufescent, longer and denser vs. almost grey, slightly shorter and sparser; head and pronotum punctures more superficial and distanced, intermediate surface almost smooth vs shagreened; eye in lateral view semilunar and more narrowed to the maxillary base and not rectilinear on frontal side, shorter to the maxilla base, on dorsal view less convex; antennomere III on dorsal view shorter than IV and slightly widened anteriorly vs. parallel; modified area on antennomeres III-VII more raised, particularly on III; pronotum sides more obliquely narrowed in front vs. more parallel.

Distribution (Fig. 8b, triangle). IRAK: Mosul; Assur (see above).

Trichomeloe sericellus (Reiche and Saulcy, 1857) (Figs 1, 5b-5d, 6f, 7a, 7c, 8c)

Meloë sericellus Reiche and Saulcy, 1857: 273, tav. 5, fig 12: Escherich, 1890: 88, 94; Reitter, 1895: 13; Jeannel and Paulian, 1944: 80, fig 28.

Meloë (Trichomeloe) sericellus, Reitter, 1911: 391; Porta, 1934: 38.

Meloë (Trichomeloe) sericellus ab. ocularis Kaszab, 1961: 531.

Trichomeloe sericellus, Bologna, 1988: 359; Bologna, 1989: 359; Bologna, 1991: 110.

Types. Reiche and Saulcy (1857) cited “Nablouse” et “Nazareth” as Type locality. Actually, we examined at the MNHN (box of “Types“) the holotype, labelled “Meloe sericellus Reiche Nazareth“ (white, handwritten), ”121 57” (white, round). Close to the holotype are positioned two additional specimens, we considered as paratypes, each only with a white, round label, respectively with the indication ”121 57“ and ”288 39“.

Diagnosis. Integument black, subopaque, head more or less extensively red, the red colouration usually is not extended on the anterior part of the frons and posteriorly to eye, laterally and ventrally and on the most part of templa; setation light yellow, dense and diffuse, short, recumbent, longer on frons, labrum and pronotum. Body length: 12.1-24 mm. Head distinctly wider than pronotum, eye small, scarcely convex, fore margin not evidently incised, anteriorly narrowed; templa 2.5 as long as the longitudinal diameter of eye, rounded, slightly wider on the middle, posteriorly very rounded; frons wide and flat, middle line almost superficial; punctures slightly deep, distanced, superficial on frons and occiput, intermediate surface shagreened. Antennomeres (Figs 5b, 7a, 7c), elongate and subcylindrical; I elongate and widened at apex, II 0.5 as long as III subglobose, III-V progressively shorter, VI-X subequal and slightly shorter than V, XI two times as long as X, parallel and conically narrowed at apex; male segments III-VII modified to form an anterior almost impunctate, shagreened area, not extended dorsally, III-VI in lateral vision evidently widened at apex. Pronotum (Figs 1, 5d) reniform, wider than long, widened in basal third and anteriorly evidently narrowed, sides rounded; base depressed and longitudinally slightly depressed in the middle; puncturation as on head; elytra vaguely rugulose. Male genitalia (Fig. 6f): in dorsal view, gonocoxal piece wide in the middle and gonostyli conically narrowed with stout apical lobes; in lateral view, gonocoxal piece wide and short, gonostyli subcylindrical on the basal third or more on ventral margin, then inclined and forming a short apical lobe; aedeagus in lateral view with two apical hooks on ventral side, similarly inclined, the prossimal one longer; endophallus hook curved.

FIG2

Fig. 7. Trichomeloe sericellus, antennomere III: a) dorsal view (the arrow indicates the modified area on anterior margin); c) detail of modified area. Trichomeloe chrysocomus, antennomere III: b) dorsal view; c) detail of anterior margin. Scale bars: a-b = 200 μm; c = 20 μm; d = 50 μm.

Remarks. Kaszab (1961) described from Sicily (see below) and Crimea, the intraspecific form ocularis, characterized by the extreme extension of the red colouration on the head. According to the Zoological Nomenclature rules, this name, described after 1960, is not available. Actually the red colouration is variable, even if usually is very extended: for example, about 80% of 13 specimens from Golan we examined, has only a semilunar red macula.

T. sericellus seems strictly related to T. deflexus and T. syriacus n.sp., based on pronotum and male genitalia shape; it differs from both species by the more extended red colouration of head, and the antennomeres IV-VII shorter than III, middle antennomeres more robust and widened anteriorly; from T. deflexus it differs also by the parameres more robust, a character in common with T. syriacus.

Distribution (Fig. 8c, spot). TURKEY: Turkey (Chikutanov, 1999). (HATAY) Akbès (MNHN; Fairmaire, 1884).

FIG2

Fig. 8. Distribution of the species of the genus Trichomeloe: (a) T. conicicollis, square; T. chyrsocomus, triangle; T. syriacus, spot; (b) T. deflexus, spot; T. mesopotamicus, triangle; (c) T. sericellus, spot; T. ottomanus, square; (d) T. ovatus, star.

SYRIA: Syria (CB; HNHM; MCSN; MHNG; MNHN; MNHU; MRT; MZR; Reiche, 1857; Baudi, 1878; Escherich, 1890b; Reitter, 1895; Pliginskji, 1911; 1914; Kaszab, 1961; Chikutanov, 1999); Golan (NHMW); Golan near El Qunetra 33°09’45”N 35°57’30”E (CB; NHMW); Krak des Chevaliers, W Homs (HNHM); Saasa (BMNH).

LEBANON: Lebanon (MNHN, MNHU); Beyrouth (MNHN); Sydon (MNHN); Ain El Zait 6 km E of El Lail (CALC); Baruk (Sahlberg, 1913).

ISRAEL and PALESTINE: Palestine (CB; HNHM; MCSN; MNHN; MNHU; Baudi, 1878; Pliginskji, 1914). Khermon-Ridge Har, Khavushit N Majdal e-Sharms (CL; CSt); Jaffa (HNHM; MNHN; NHMW); Mt. Carmel (MNHN); Haifa (HNHM, MNHN); Tel Aviv (CB); Lake Tiberias, Genezareth (Sahlberg, 1913); Tel Aviv, Ben Shemen (CB; MZUR); Galilaea (MNHU); Galilaea, Mt. Meron (CB); Galilaea, Mt. Meron, Ya’ar’am (CL; CSt); Nazrath (MCSN; MNHN; SMNS; Reiche and Saulcy, 1857; Kaszab, 1961); Ain Zabassim (MUH); Nablous (Reiche and Saulcy, 1857; Borchmann, 1917); Bethlem (MNHN); Jerusalem (MHNG; HNHM; MSNM; MSNV; MNHN; MNHU; Sahlberg, 1913); Kiryat Anavim (Bodenheimer, 1934); Kubbet Râhil (Sahlberg, 1903); Der Aban (Sahlberg, 1913); near Birel Maksur (CL; CSt); West Bank, Jerusalem (HNHM; MHNG); Negev, South of Sede Boqer (CL; CSt). Chikatunov (1999) cited the species from the following areas: Upper Galilee, Lower Galilee, Golan Heinghts, Mount Hermon, Samaria, Valley of Yzre’el, Judean Hills, Dead Sea, central coastal plain, northern Negev.

Uncertain locality: Sarvid (JP).

Some uncertain records were quoted in the literature. In particular, this species was cited from Palermo (Luigioni, 1929; see also Bologna, 1991) and generically from Sicily (Porta, 1934; Kaszab, 1961; see also Bologna, 1991, 1995). At the HNHM is preserved one specimen (paratype of the invalid form ocularis), we examined, labelled “Sicilia” and studied by Kaszab (1961). The citation from Naples (as “Napoletano”: Luigioni, 1929, or as “Campania”: Porta, 1934), could derive from an error made by Borchman (1917), who indicated “Neapel”, as misinterpretation of “Nablous”, a Palestinian town. Alternately, if the presence in Sicily is confirmed, the Luigioni’s citation could indicate the old name of the Naples Kingdom, which included Sicily until 1861. Kaszab (1961), on the base of one specimen (paratype of the unvalid form ocularis) quoted T. sericellus also from Crimea (southern Ukraina).

The species, as the most part of the genus, seems to be endemic to the Levant and Middle East, and both the eccentric records (Sicily and Ukraina) seem to be rejected.

Trichomeloe syriacus Bologna n. sp. (Figs 6d, 8a)

Type material. 1 male holotype from “Sy. Tadmur, 15.iii.1980, Gräber von Palmyra, R. Kinzelbach leg.” (CB). 1 female paratype from “Syria, Palmyra, 30.i.1989, P. De Salvo leg.” (CB). 1 male and 1 female paratypes from “Syrie, Rankous, 08.iii.02, P. Weill” (CB and CW).

One female paratype has antennomeres slightly damaged.

Other specimens examined. 1 specimen from Palmyra, found dead and very damaged (without appendices), was not considered as paratype. Other specimens from Palmyra, previously identified as T. deflexus, are preserved at SMNS, but were not re-examined during this revision.

Description. Integument shiny black, with one small red and suboval frontal spot; setation very short and black, sparse on head, pronotum and elytra, ventrally light grey longer and quite dense. Body length: 14.0-22.1 mm.

Head wider than pronotum, eye small and convex, in lateral view fore margin slightly incised, short and slightly narrowed anteriorly; templa 2.5 as long as the longitudinal diameter of eye, sub-parallel, slightly wider and rounded posteriorly; middle frontal line superficial, but well visible; punctures deep but spaced, intermediate surface shagreened. Antennomeres robust, elongate and subcylindrical; I elongate and widened at apex, II ca 0.5 as long as III subglobose, III antennomere slightly shorter or subequal to IV; VI-X subequal and similar in length to V, XI twice as long as X, parallel and conically narrowed at apex; male segments III-VII modified to form an anterior almost impunctate, shagreened area, slightly extended also dorsally, III-VI in lateral vision not distinctly widened at apex. Clypeus short and transverse; labrum transverse, short, emarginate anteriorly; both with puntures similarly than on head capsule.

Pronotum short, subreniform, more than twice as wide as long, sides subrounded from base; base evidently depressed, sides dorsally evidently convex (as in some species of the Meloe brevicollis complex); middle surface longitudinally slightly depressed; punctures and surface as on head, but slightly denser. Prosternum short and sub-transverse. Elytra vaguely rugulose, sub-alutaceous. Legs robust, with robust black setae.

Abdominal tergites with fine and scattered punctures and quite sparse setae; posterior margin of both last male tergite and sternite with long setae. Male genitalia (Fig. 6d): in dorsal view, gonocoxal piece short, wide in the middle, gonostyli robust, conically narrowed with robust and stout apical lobes; in lateral view, gonostyli robust, apical lobes elongate but robust, with long setae on both sides; aedeagus in lateral view with two apical hooks on ventral side, both short, the prossimal one great and apical in position.

Remarks. Among the species related to T. sericellus, this new species seems close particularly to T. sericellus and T. deflexus because it lacks modified body setation and the integument colour. It differs from T. sericellus, which has similar male gonostyli, by the reduced red frontal colouration, the shape of male middle antennomeres not widened apically, the shape of eye. Before this revision some specimens of the new species were identified as T. deflexus also by one of us (M.A.B.). Actually T. syriacus differs from T. deflexus because the head and pronotum setae black, shorter and more robust, as well as by the parameres more robust, particularly the apical lobes, and the shape of aedeagal hooks, which are more apical in position. Escherich (1890) consider as T. sericellus one variety with small red colouration, which could correspond to T. syriacus.

Distribution (Fig. 8a, spot). SYRIA: Palmyra; Rankous (see above)

Subgroup B

Trichomeloe ottomanus (Pliginskji, 1914) (Figs 6g, 8c)

Meloë ottomana Pliginskji, 1914: 254.

Meloë (Trichomeloe) frivaldszky [sic] Kaszab, 1958: 304.

Trichomeloe ottomanus, Bologna, 1988: 30; Bologna, 1989: 359; Bologna, 1991: 110.

Type material. Types of this species are preserved in the Motschulsky’s collection, probably at the Moscow Museum. The type locality of the species is “Armain” (Pliginskji, 1914), a late name indicating most of the eastern Turkey and the Armenia Republic.

The holotype and 1 paratype of the synonym frivaldzky from South-East Turkey (Elazig), we examined, are preserved at the HNHM.

Diagnosis. Integument shiny black, but head reddish, except clypeus, in some specimens a small frontal spot, a short oval spot extended from the posterior margin of eye, the ventral side black; pronotum vaguely brownish on sides; setation light yellow, dense and diffuse, short, recumbent, longer on frons, labrum and pronotum; pronotum and abdomen with elongate light setae, distributed on tufts on abdomen, separated by nude areas. Body length: 11-22 mm. Head with a short and shallow depression posteriorly to eye, evidently wider than pronotum; eye small, scarcely convex; templa more than 2.5 as long as the longitudinal diameter of eye, wider on the middle, posteriorly rounded; frons wide and slightly convex, middle line almost superficial; punctures wide, deep, distanced, intermediate surface shagreened, frons medially almost unpunctate. Mandibles dorsally depressed, wide. Antennomeres, elongate and subcylindrical; I not evidently elongate, II 0.5 as long as III subglobose, III as long as V and slightly longer than IV, VI-X subequal and slightly shorter than V, XI two times as long as X, parallel and conically narrowed at apex; male segments III-VII modified to form an anterior almost impunctate, shagreened area, slightly extended dorsally. Pronotum subpentagonal with smooth and nude raised areas on sides and longitudinally in the middle; base depressed and longitudinally evidently and largely depressed in the middle; puncturation as on head; elytra densely and finely rugulose, subopaque. Abdominal tergites densely setose but with shiny glabrous spots on sides and longitudinally in the middle; sternites III-V with anterolateral unsetated spotted surface. Male genitalia (Fig. 6g): in dorsal view, gonocoxal piece wide in fore half and gonostyli conically narrowed with wide and stout apical lobes; in lateral view, gonocoxal piece wide and short, gonostyli subcylindrical on the basal third or curved dorsally in the middle, then inclined to form a short and very stout apical lobe; aedeagus in lateral view with two apical hooks on ventral side, similarly inclined, the proximal one longer.

Remarks. This species differs considerably phenetically from others of the genus Trichomeloe because of the head punctures and colouration, mandible shape, the less reniform pronotum and anteriorly more angulate, medially depressed with raised nude areas, the abdomen densely setose. Interestingly, it shows several convergences in the integumentary structure with Meloe (Lasiomeloe) olivieri Chevrolat, 1833, endemic to Turkey.

Distribution (Fig. 8c, square). TURKEY: Anatolia (Kaszab, 1958). Armenia (Pliginskji, 1914). (ELAZIG) Elazig (HNHM).

SYRIA: Syria (JP; MNHN). Possibly this indefinite locality indicates late regions of the Ottoman Empire, now in southern Turkey.

Some specimens without locality label were also examined (CB; MNHN).

Trichomeloe ovatus (Pripisnova, 1987) (Figs 8d)

Meloë (Trichomeloe) ovatus Ppipisnova 1987: 43.

Trichomeloe ovatus, Bologna and Pinto, 2002: 2055.

Type material. According to the description, the male holotype, is preserved in the Tajikistan Academy of Sciences, and 1 male paratype at the San Petersberg Museum. We requested several times these specimens for study, but we did not obtain answer from the first institution, and the paratype was not found by the Curator in the second Museum.

The Type locality, Tigrovaja balka, is a Natural Reserve in the Amudarja region, about 180 km S of Dushambe, close to the border with Afghanistan, characterized by different ecosystems, from desert to forest and marshes. This species probably was collected in semidesert or steppe habitats.

Diagnosis (from the description). Body leaden black, shiny; setation light yellow, dense and diffuse, short, recumbent, disposed almost on tufts on head sides, uniformly and densely distributed on pronotum and elytra; abdomen with light setae, spaced out by nude areas. Body length 7-10 mm. Head subquadrate, frons convex, with a middle longitudinal line and two small projections overlooking the clypeus, base evidently emarginated at base; eye convex; punctures wide, subrugose on frons. Antennomeres elongate and subcylindrical, I-IV shiny, V-XI opaque. Pronotum slightly wider than head, subtrapeizoidal, with fore angles rounded and with one rounded depression, posterior angles less rounded, base depressed, longitudinally deeply and largely depressed in the middle; punctures wide; elytra densely and finely rugulose, subopaque, humeri slightly bulging. Abdominal tergites densely setose but with one shiny glabrous spot on each side; sternites glabrous and smooth. Male genitalia: in dorsal view, gonocoxal piece wide in the middle, gonostyli conically narrowed with slender apical lobes; in lateral view, gonocoxal piece wide and short, gonostyli subcylindrical on the basal third or more widened in the middle on ventral margin, then inclined to form a short and stout apical lobe; aedeagus in lateral view with two distanced apical hooks on ventral side, similarly inclined; endophallus hook curved.

Remarks. Species almost unknown. According to the description it seems to have some similarity with T. ottomanus. For this reason, we tentatively assigned it to a subgroup with the Turkish species.

Distribution (Fig. 8d, star). TAJIKISTAN. Tigrovaja balka (Pripisnova, 1987).