Contributions to Zoology, 86 (1) – 2017Isabel T Hyman; Irantzu de la Iglesia Lamborena; Frank Köhler: Molecular phylogenetics and systematic revision of the south-eastern Australian Helicarionidae (Gastropoda, Stylommatophora)

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Peloparion Iredale, 1937

Peloparion Iredale, 1937a: 8.

Type species: Helicarion helenae Godwin-Austen, 1883, by subsequent designation of Iredale (1941); masculine


Diagnosis. Shell. Small, ear-shaped, flattened, last whorl large, umbilicus closed, shell glossy, protoconch with fine spiral grooves, teleoconch smooth (Table 3, Fig. 35).


Fig. 35. Shells of Peloparion and Ubqiuitarion. A. Peloparion helenae: holotype of Peloparion submissus Iredale, 1941, AM C.101142 (size indicative; photographer: AM). B. Ubiquitarion iridis holotype AM C.456552. Scale bar = 3 mm.

Animal. Pale grey with bright red mucus. Mantle lobes and shell lappets well developed.

Genitalia. Ovotestis of four lobes, embedded in digestive gland. Talon and carrefour embedded in albumen gland. Spermoviduct folded twice. Free oviduct very long with no internal sculpture; capsular gland not externally visible. Bursa copulatrix moderately short, reaching less than halfway along spermoviduct; internally smooth; inserted on vagina. Vagina very short. Epiphallus enters penis through moderately long verge; epiphallic caecum absent; epiphallic flagellum with axial filament present; numerous internal cryptae present in part of epiphallus adjacent to flagellum but not in flagellum. Spermatophore a soft-walled capsule with hard tail-pipe; branching spines present on capsule; tail-pipe with only a single spine.

Remarks. Peloparion was originally introduced as a subgenus of Helicarion for Vitrina hyalina Pfeiffer, 1855 (Iredale, 1937c), but subsequently Iredale (1941) stated that the name was intended for Helicarion helenae Godwin-Austen, 1883. The genus was revised by Hyman (2007) and Hyman and Ponder (2010).