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Contributions to Zoology, 67 (1) 3-8 (1997)


Jan Hendrik Stock, 22 February 1931 - 17 February 1997

Live and Career of Jan Stock


Jan H Stock (Photo: Louis van der Laan)

Jan Stock was born in western Amsterdam 22 February 1931, the only child of Jan Hendrik Stock, a bank clerk, and Elisabeth Stock-Knevel. He exhibited an early interest as a child in living things, raising plants on a back balcony of his parents apartment and collecting animals during family excursions to the beaches at Zandvoort. However, it was in his early teens in the Hoogere Burgerschool that his interest in biology blossomed where he achieved perfect scores in natural history on his final examinations. He entered the University of Amsterdam in 1948 to pursue his studies of biology and worked under such figures as Profs. L.F. de Beaufort and H. Engel - his predecessors in the chair of Special Zoology - as well as Profs. J.E.W. Ihle and E.J. Slijper. It was Ihle in fact who first encouraged Stock to take up the study of pycnogonids, the sea spiders, a group in which he made some of his initial scientific contributions.

His first scientific articles date from the student years and appeared in Het Zeepaard, the journal of the Dutch Youth-Union for Nature Studies and the Dutch Natural History Society. These attracted him to the attention of De Beaufort who gave him a job as a curatorial assistant in the Zoological Museum of Amsterdam in 1950 with duties in the then poorly organized collections of marine invertebrates. He set up a card catalogue of the collections, tracked loans of museum materials to other researchers, and identified specimens for members of the public. It was through this last activity, when he received from the Mussel Institute at Bergen-op-Zoom in the Dutch province of Zeeland some parasitic copepods afflicting mussels, that Stock developed another of his research interests, the taxonomy of copepods. His first real scientific papers date from that year.

Stock completed his university degree in 1955, graduating cum laude, and in record time he defended his Ph.D. thesis under Engel, "The pycnogonid family Austrodecidae," in 1957. The comments solicited from three foreign members of his examining committee, Isabella Gordon of the British Museum, Joel Hedgpeth of the old Pacific Marine Station in California, and Ernst Mayr of Harvard, helped to insure that Stock received a cum laude for that effort as well.

For the next 11 years he produced a massive amount of research work, most of it in the systematics of pycnogonids and parasitic copepods, much of the latter in French dealing with the parasitic forms from the coasts of France. Stock‘s first publications concerning gammarid amphipods, for which he was so well known later, appeared in only 1966; and his interests and publications soon ranged across the Eurasian landmass - from Holland to Lake Baikal.

He married Brenda Balfoort in 1964 and they had four sons. In 1968 with the retirement of Engel, Stock succeeded to the chair. Among his duties as professor was to act as editor of this journal (then known by its Dutch name as Bijdragen tot de Dierkunde). He was also instrumental in establishing the Bulletin Zoologisch Museum Amsterdam for the publication of purely taxonomic studies. It was about this time that his full fledged interest in ground water faunas began to develop, although it was not until 1976 that his initial scientific publications, in thermosbaenaceans and ingolfiellid amphipods, actually appeared. From this point on, Stock‘s publications came to increasing deal with various ground water crustaceans, in addition to those just mentioned also including bogidiellid and hadziid amphipods, isopods, and various free-living copepods. Indeed his work on ground water faunas led to his helping to found the journal Stygologia, which quickly became an important journal in the field and remained so for years but which unfortunately was eventually discontinued by the publisher for financial reasons. From 1980 onward he concentrated on supposed Tethys derived biotas in the Caribbean, the tropical and subtropical Atlantic islands and the peri-Mediterranean belt. With unrelenting stamina he sampled these islands and nearby coastal areas, and those who accompanied him will remember the experienced way in which he tackled all the practical problems - from getting permits from suspicious landowners to giving lectures in every conceivable circumstance. From his extensive collections he extracted a general theory, which he called the Regression Model. This is an essentially vicariant scenario that tries to explain and predict how several Tethyan, marine ancestors became adapted to groundwater conditions during periods of relative low sea level, and how these descendants are found and will be found in those areas that are uplifted.

By the time he retired, Jan Stock had published some 448 articles. Freed of his academic responsibilities, retirement only increased his productivity. At the time of his death he had published more than 55 additional papers, and several items are now in press or ready to be submitted - a truly astounding level of productivity of around 11 papers per year.

An intensely private man, he good naturedly erected a screen of fabulous stories around his life. For example, one of us (FRS), quite believed that Stock was born in the Dutch East Indies because Jan had most convincingly told him so many years ago, and it was only in preparing this obituary that he was surprised to learn that Stock was born right in Amsterdam. Stock‘s humor was legendary and he revelled in regaling students and colleagues with some of the most elaborate and tongue-in-cheek tales. "A good story is a good story, no matter if it is true or not," he once said.

He will be remembered in the University of Amsterdam for his interests in teaching, especially in field situations, and for his service in biology faculty administration, having served as dean for some years. His intense enjoyment of field work resulted in, among other trips, major expeditions to the Red Sea, various Atlantic Islands, and participation in the Dollard-Eems project in the Wadden Sea. This devotion to field work continued after his retirement, and he organized another two major expeditions in the years between his retirement and death.

A workaholic himself, Stock nevertheless left his students and colleagues alone to find their own way. He would gladly give advice when asked, and he promptly read and returned manuscripts when they were given to him. However, he firmly believed that everyone should find their own level of productivity. Jan Stock will be greatly missed around the Zoological Museum of Amsterdam, but his scientific work in the systematics of marine arthropods is an enduring monument. A bibliography of Jan Stock up to 1990 can be found in Weinberg & van Zijl (1990, Bulletin Zoölogisch Museum, Spec. Vol. 1990:1-42) and an update of his publications published since then is included here.

Additional publications of J.H. Stock

Stock, J.H., 1990. Review of Bruce E. Felgenhauer, Les Watling & Anne B. Thistle (eds.) Functional morphology of feeding and grooming in Crustacea. Crustacean Issues, 6, 1989. Crustaceana, 59: 319.

Stock, J.H., 1990. Review of S. Ruffo (ed.) The Amphipoda of the Mediterranean, part 2 (Mémoires de l‘Institut océanographique, Monaco, no. 13, 1989). Crustaceana, 59: 319.

Stock, J.H., 1990. Insular groundwater biotas in the (sub) tropical Atlantic: A biogeographic synthesis. Atti dei conv. Accad. Naz. Lincei (Roma), 85: 695-713.

Stock, J.H., 1990. A new species of Austrodecus (Pycnogonida) from New South Wales, Australia. Tijdschr. Entom, 133: 269-272, figs 1-9.

Stock, J.H. & E. Sánchez, 1990. First record of Crustacea Malacostraca from fresh waters in the Canary Islands. Hydrobiologia, 206: 53-59.

Stock, J.H. & R. Vonk, 1990. A freshwater amphipod from La Gomera, Melita dulcicola n.sp. (Stygofauna of the Canary Islands, 23). Annls. Limnol., 26: 29-37.

Stock, J.H., 1990. Sur deux Pycnogonides armoricains remarquables. Cah. Biol. mar., 31: 513-516.

Stock, J.H., 1991. Some reflections on the antiquity of the copepod lineages. Proc. Fourth Int. Conf. Copepoda; Bull. Plankton Soc. Japan, Spec. Vol. (1991): 1-7.

Stock, J.H., 1991. Pycnogonides de la campagne Seamount 1 au large de la péninsule ibérique et dans le golfe inéro-marocain. Bull. Mus. natl. Hist. nat. Paris, 4e sér., 13, section A: 135-142.

Stock, J.H. & T.M. Iliffe, 1991. Two new species of Liagoceradocus (hypogean Amphipoda) from south-western Pacific Islands, with key to the world species. Invertebr. Taxon., 5: 807-25.

Stock, J.H., 1991. A new species of Psammogammarus (Amphipoda, Melitidae) from river alluvia in Luzon, Philippines. Stygologia, 6: 227-233.

Stock, J.H. & R. Vonk, 1991. Une espèce nouvelle de Dulzura, genre d‘Amphipodes hadzioïdes connu jusqu‘ici seulement de l‘Indo-Pacifique, découverte aux îles du Cap-Vert (océan Atlantique). Cah. Biol. Mar., 32: 477-486.

Stock, J.H, 1991. Deep-water Pycnogonida from the surroundings of New Calidonia. In: A. Crosniers (ed.), Résult. des Camp. MUSORSTOM, 8., Mém. natn. Hist. nat., (A) 151: 125-212.

Stock, J.H, 1991. Pycnogonida of the MUSORSTOM campaigns to the Philippines. In: A. Crosnier (ed.) Résult. des Campagnes MUSORSTOM, volume 8. Mém. Mus. natn. Hist. nat. (A), 151: 213-228

Vonk, R. & J.H. Stock, 1991. Caecostenetroides ascensionis n. sp., a blind marine interstitial isopod (Asellota, Gnathostenetroidoidea) from Ascension Island, South Atlantic. Trop. Zool., 4: 89-98.

Pretus, J.L. & J.H. Stock, 1991. A new hyporheic Bogidiella (Crustacea, Amphipoda) from Mallorca. Endins, no. 16: 47-51.

Humes, A.G. & J.H. Stock, 1991. Coralliomyzontidae, fam. n. (Copepoda: Siphonoistomatoida) associated with scleractinian corals in Madagascar. Bull. Zool. Mus. Univ. Amsterdam, 13: 17-24.

Stock, J.H. & D. Platvoet, 1991. The freshwater Amphipoda of the Falkland Islands. J. Nat. History, 25: 1469-1491.

Stock, J.H., 1992. Bogidiella (Amphipoda) in Japanese inland waters. Crustaceana, 62: 273-282.

Stock, J.H., 1992. Pycnogonida from southern Brazil. Tijdschr. Entom., 135: 113-139.

Stock, J.H., 1992. Shifting ranges and biodiversity in animal ecosystems. In: O.T. Solbrig, H.M. van Emden & P.G.W.J. van Oordt, Biodiversity and Global Change. Intern Union Biol. Sci., Monogr. 8: 167-171.

Stock, J.H. & R. Vonk, 1992. Marine interstitial Amphipoda and Isopoda (Crustacea) from Santiago, Cape Verde Islands. Bijdr. Dierk., 62: 21-36.

Stock, J.H. & R. Vonk, 1992. The first freshwater amphipod (Crustacea) from the Cape Verde Islands: Melita cognata n.sp., with notes on its evolutionary scenario. Revue Zool. Afr., 106: 273-280.

Stock, J.H., 1992. Discovery of the male and subgeneric status of Bogidiella cypria Karaman, 1989 (Crustacea, Amphipoda). Stygologia, 7: 117-118. 411.

Stock, J.H., 1992. Littoral Pycnogonida from Oman. Bijdr. Dierk., 62: 81-98.

Stock, J.H., 1992. Bogidiella (Amphipoda) in Japanese inland waters. Crustaceana, 62: 273-282.

Stock, J.H., 1993. Pycnogonida: Description d‘Ascorhynchus miniscapus sp. nov., récolté sur le banc de la Bayonnaise (nord-ouest des îles Wallis et Futuna). Résultats des campagnes MUSORSTOM, 11. Mém. Mus. natn. Hist. nat., 158: 349-353.

Stock, J.H., 1993. Some remarkable distribution patterns in stygobiont Amphipoda. J. Nat. History, 27: 807-819.

Stock, J.H., 1993. Copepoda (Crustacea) associated with commercial and non-commercial Bivalvia in the East Scheldt, The Netherlands. Bijdr. Dierk., 63: 61-64.

Stock, J.H., 1993. How endangered is Gammarus duebeni celticus, a native freshwater amphipod in western Brittany (France). Arch. Hydrobiol., 128: 415-421.

Stock, J.H., & A.D. Abreu, 1993. Three new species of Pseudoniphargus (Crustacea: Amphipoda) from the Madeira Archipelago. Bol. Mus. Mun. Funchal, 44: 131-155.

Stock, J.H., 1994. De "slijkgarnaal" Corophium sextonae (Amphipoda) plotseling talrijk in de Oosterschelde. Het Zeepaard, 54: 82-84

Stock, J.H., 1994. Biogeographic synthesis of the insular groundwater faunas of the (sub)tropical Atlantic. Hydrobiologia, 28: 105-117.

Stock, J.H. & C.K. Biernbaum. 1994. Terrestrial Amphipoda (Talitridae) from Ascension and Saint Helena (South Central Atlantic). J. Nat. Hist., 28: 795-811.

Stock, J.H., 1994. Shifting ranges and biodiversity in animal ecosystems. In: O.T. Solbridge, H.M. van Emden & P.G.W.J. van Oordt, Biodiversity and Global change. CAB International, 1994: 169-173.

Beyer, G. & J.H. Stock, 1994. Epigean freshwater gammaridae (Crustacea, Amphipoda) from la Gomera (Canary Islands). Bijdr. Dierk., 64: 101-114.

Stock, J.H., 1995. Vindplaatsen van de Ivoorpok, Balanus eburneus, in Nederland. Het Zeepaard, 55: 19-22.

Stock, J.H., 1995. Two new poecilostomatoid copepods (Crustacea) associated with the polychaeta Spirobranchus from the Seychelles. J. Afr. Zool., 209: 77-88.

Stock, J.H., 1995. De eerste vondsten van het zeepokken-geslacht Chthamalus in Nederland. Het Zeepaard, 55:119-123.

Stock, J.H., 1995. Orchestia aestuarensis (Amphipoda, Talitridae) nu ook in Nederland aangetroffen. Het Zeepaard, 55:81-84.

Stock, J.H., 1995. De glasvlooien (Amphipoda, geslacht Hyale) van Nederland. Het Zeepaard, 55: 46-47.

Stock, J.H., 1995. The genus Sarothrogammarus martynov, (Crustacea, Amphipoda) in Madeira. Biol. Mus. Mun. Funchal, 45: 29-40.

Stock, J.H., 1995. A new member of the family Bogidiellidae (Crustacea, Amphipoda from poikilohaline waters of Madeira. Bocagiana, 171: 1-8.

Stock, J.H., 1995. Gammarus and Cheatogammarus (Crustacea, Amphipoda) from Madagascar. Bol. Mus. Mun. Funchal, 247: 41-52.

Stock, J.H., 1995. Copepoda Poecilostomatoida associated with Bivalvia from new Guinea. Hydrobiologia, 312: 37-45.

Stock, J.H., 1995. Two new poecilosytomatoid copepods (Crustacea) associated with the polychaete Spirobranchus from the Seychelles. J. Afr. Zool. 109: 77-88.

Stock, J.H. 1995. Origin and evolution of the island of Saint Helena (South Atlantic). List of sampling stations April-May 1995. Verslagen en Technische gegevens, Zool. Mus. Amsterdam, 63: 1-9.

Miyazaki, K. & J.H. Stock, 1995. Rhynchothorax mediterraneus, the first record of Rhynchothoracidae (Pycnogonida) from Japan. Publ. Seto Marine Biol. Lab., 36: 325-327.

Stock, J.H., 1996. Sabelliphilus elongatus, een parasitaire Copepode van de kokerworm Sabella, in de Oosterschelde. Het Zeepaard, 56: 14-15.

Stock, J.H., 1996 (1995). Biogeography and evolutionary scenario of aquatic organisms in Macaronesia. Bol. Mus. Mun. Funchal, 4: 729-745.

Stock, J.H. & J.C. von Vaupel Klein. 1996. Mounting media revisited: the suitability of Reyne‘s fluid for small Crustaceans. Crustaceana, 69: 793-798,

Stock, J.H., 1996. Age of limnic and stygobiont Amphipoda of the Madeira Archipelago and La Gomera, Canary Islands. Polsk. Arch. Hydrobiol., 42: 409-415.

Stock, J.H., 1996. The genus Platorchestia (Crustacea, Amphipoda) on the Mid-Atlantic islands, with description of a new species from Saint Helena. Miscel. Zool., 19: 149-157.

Stock, J. H., 1996. Two new species of Copepoda parasitic on polynoid polychaetes. Crustaceana, 69: 438-445.

Kim, Il-Hoi & J.H. Stock, 1996. A new species of Clausidiidae (Copepoda, Poecilostomatoida) associated with the bivalve Ruditapes philippinarum in Korea. Cah. Biol. Mar., 37: 1-6.